Celastrina argiolus (Linnaeus, 1758)
NAF, EU, Siberia, Turkey, C.Asia, Japan. See [maps]
Tosteblåvinge
Holly Blue
Najade
Paatsamasinisiipi
Skogsblåvinge
Modraszek wieszczek
Faulbaumfalter
- Papilio argiolus Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 483; TL: England
- Papilio cleobis Sulzer, 1776; Gesch. Ins. nach linn. Syst. (2): pl. 18, f. 13-14
- Papilio argyphontes Bergsträsser, [1779]; Nomen. Ins. 2: 15, pl. 58, f. 5-6
- Papilio argalus Bergsträsser, [1779]; Nomen. Ins. 2: 18, pl. 60, f. 4-5
- Papilio thersanon Bergsträsser, [1779]; Nomen. Ins. 3: 4, pl. 49, f. 5-6
- Papilio marginatus Retzius, 1783; Gen. Spec. Ins.: 30
- Papilio acis Fabricius, 1787; Mantissa Insectorum 2: 73, n. 687
- Plebeius argiolus var. paraleuca Röber, 1897; Ent. Nachr. 23 (17-18): 270
- Celastrina argiolus ab. c-nigrum Tutt, [1908]; Nat. Hist. Br. Lepid. 9: 398
- Celastrina argiolus var. kobei Tutt, [1908]; Nat. Hist. Br. Lepid. 9: 401
- Cyaniris argiolus ab. aquilina Grund, 1908; Int. ent. Zs. 2 (14): 88
- substusradiata (Blachier, 1909) (Cyaniris); Bull. Soc. Lep. Genève 1: 381
- Cyaniris argiolus f. aest. latisquama Ball, 1914; Ann. Soc. ent. Belg. 58: 180, pl. 4, f. 21-22
- Lycaenopsis argiolus r. britanna Verity, 1919; Ent. Rec. J. Var. 31: 46
- Lycaenopsis argiolus f. calidogenita Verity, 1919; Ent. Rec. J. Var. 31: 46
- Lycaenopsis argiolus f. mixta Verity, 1919; Ent. Rec. J. Var. 31: 47
- microdes (Cabeau, 1925) (Cyaniris); Rev. mens. Soc. ent. Namur. 25: 7
- anteatrata (Mezger, 1934) (Lycaenopsis); Lambillionea 34: 151
- expansa (Pionneau, 1937) (Cyaniris); Echange 53: 3; TL: France
- Celastrina argiolus f. ♀ antidisconulla Lempke, 1954; Tijdschr. Ent. 97: 344
- Celastrina argiolus f. ♀ maculispostcarens Lempke, 1954; Tijdschr. Ent. 97: 345
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- Celastrina (Celastrina) argiolus; [RFEL]
- Celastrina argiola; [MRS], 680
- Celastrina argiolus antidisconulla; [CoL]
- Celastrina argiolus britanna; [CoL]
- Celastrina argiolus calidogenita; [CoL]
- Celastrina argiolus maculispostcarens; [CoL]
- Celastrina argiolus; Forster, 1941, Mitt. münchn. ent. Ges. 31 (2): 613; [SPRK]; [H&R]; [BARU], #358; [EBW]; [BOW]: pl. 9, f. 13-14; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BRU2]: 143, pl. 63, f. 16-27, 31-36; [NL4A], #1086; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 79; Yakovlev, 2012, Nota lepid. 35 (1): 71; [FE]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [SVF] ; [Mario Maier]
- Celastrina ladon argiolus; [CoL]
- Cyaniris argiolus; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 304
- Lycaena argiolus; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 414
- Lycaenopsis argiolus; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 443; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 27; [iNaturalist]
500x605 (~31Kb) underside Finland, Tampere, Hervanta, 681:33, 12.5.1996, Photo © Tero Piirainen
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533x400 (~41Kb) ♂ underside A road at a birch grove edge near the village Mel'nichikha (Shadrikha), Novosibirsk District and Province, West Siberia, Russia. 18th May 1997, Photo © Oleg Kosterin
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700x556 (~87Kb) ♂ underside The Inya River right bank at Razyezd Inya station, Novosibirsk, West Siberia, Russia. 22nd April 2000, Photo © Oleg Kosterin
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1000x1186 (~160Kb) underside Russia, Udmurtia, river banks of Kama, near village Doksha east of Izhevsk, 20.8.2004, Photo © Markku Savela
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800x600 (~115Kb) underside Russia, Kaluga area, 19.05.2005, Photo © D. Smirnov
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701x851 (~121Kb) ♂ Russia: Kaluga area (35°45'E, 54°23'N), 19.5.2005, Photo © D. Smirnov
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500x417 (~56Kb) Turunc, SW Turkey 19/05/92, Photo © Simon Coombes
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Larva on
(as argiola) Sophora flavescens,
Lespedeza spp.,
Desmodium oxyphyllum,
Vicia sativa,
Pueraria lobata,
Wisteria floribunda,
Cornus controversa,
Aralia elata,
Euodia meliifolia,
Malus pumila,
Prunus salicina,
Polygonum cuspidatum,
Quercus aliena,
Staphylea bumalda [MRS],
Ribes ,
R. nigrum,
R. rubrum,
R. uva-crispa,
Rhamnus frangula,
R. cathartica [SPRK],
Caragana frutex,
Chamaecystis ,
Lupinus ,
Vaccinium [BARU],
Hedera helix,
Ilex aquifolium,
Genusta florida Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21: 36,
Berberis ,
Cuscuta engelmannii,
C. lehmanniana,
(C. Asia) [BRU2]: 144
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C. a. argiolus EU, Caucasus, Armenia, Talysh Mts., Kazakhstan, Atai, Sayan, W.Siberia, C.Siberia
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C. a. parvipuncta (Fuchs, 1880)
- Celastrina argiolus parvipuncta; [CoL]
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C. a. mauretanica (Rothschild, 1925) Morocco, Algeria, Tunisia
- Celastrina argiolus mauretanica; [BMAT]: 35, pl. 11, f. 25-30; [CoL]
Larva on
Arbutus unedo,
Rubus sp.,
Discolores sp.,
Cornus sp.,
Rhamnus sp.,
Colutea sp. [BMAT]
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C. a. bieneri Forster, 1941 Yakutia, Transbaikalia, Far East, Kamchatka
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C. a. caphis (Fruhstorfer, 1922) Yunnan, ...?
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C. a. sikkima (Moore, [1884]) Sikkim - Assam, Karens
Hill Hedge Blue
Larva on
Prinsepia utilis [BIR]
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?C. a. kollari (Westwood, [1852]) Chitral - Kumaon
Hill Hedge Blue
- Celastrina argiolus kollari; [iNaturalist]
- Cyaniris coelestina kasmira; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 294
- Cyaniris coelestina; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 294
- Lycaena kollari; [CoL]
- Lycaenopsis argiolus coelestina; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 29
- Lycaenopsis argiolus coelestina f. coelestina; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 29
- Lycaenopsis argiolus kollari; [BIR], 279(key), 280
Larva on
Prinsepia utilis [BIR]
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C. a. iynteana (de Nicéville, 1884) Nepal - Burma, Thailand, Vietnam, Tibet, S.Yunnan
Jyntea Hedge Blue
- Cyaniris iynteana de Nicéville, [1884]; J. Asiat. Soc. Bengal 52 Pt.II (2/4): 69, pl. 9, f. 7 ♂, 7a ♀; TL: Sikkim
- Cyaniris jynteana Moore, [1884]; Proc. zool. Soc. Lond. 1883 (4): 524, pl. 48, f. 10; TL: Khasia and Jyntea Hills
- Cyaniris victoria Swinhoe, 1893; Trans. ent. Soc. Lond. 1893 (3): 293; TL: Shillong, Assam
- Lycaenopsis argiolus var. bothrinoides Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 446
- Lycaenopsis argiolus var. puspargiolus Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 446; TL: Assam
- Celastrina argiolus var. albocaeruleoides Tutt, [1908]; Nat. Hist. Br. Lepid. 9: 404 (Chapman in litt.)
- Lycaenopsis argiolus var. albocaeruleoides Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 446
- Cyaniris jynteana herophilus Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 293; TL: Vietnam
- Cyaniris coelestina cition Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 294; TL: Vietnam
- Celastrina albocoeruloides; [CoL]
- Celastrina argiolus albocaeruleoides; [CoL]
- Celastrina argiolus iynteana; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 106 (note); [Yutaka Inayoshi]
- Celastrina iynteana herophilus; [CoL]
- Celastrina iynteana; [CoL]
- Celastrina jynteana; Forster, 1941, Mitt. münchn. ent. Ges. 31 (2): 611
- Celastrina puspargiolus; [CoL]
- Cyaniris jynteana; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 293
- Cyaniris victoria; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 283, 303; [CoL]
- Lycaenopsis argiolus herophilus; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 29
- Lycaenopsis argiolus victoriae[sic, victoria]; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 29
- Lycaenopsis jynteana; [BIR], 280
- Lysandra coelestina cition; [CoL]
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C. a. hypoleuca (Kollar, [1849]) Kopet-Dagh, Tian-Shan, Ghissar-Darvaz, Pamirs-Alai
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C. a. ladonides (d'Orza, 1869) Amur, Ussuri, Sakhalin, Kuriles, Japan
Silvery Hedge Blue
- Lycaena ladonides d'Orza, 1869; Lep. Jap.: 20
- Lycaena levettii Butler, 1883; Ann. Mag. nat. Hist. (5) 11 (62): 111
- Celestrina[sic] (Cyaniris) sachalinensis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 105, pl. 8, f. 16 ♂ (preocc.); TL: Sakhalin
- Cyaniris argiolus (gen. aest.) heringi Kardakoff, 1928; Ent. Mitt. 17 (4): 272, pl. 7, f. 20-21; TL: Primorye
- Lycaenopsis manchurica Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 358; TL: Manchuria
- Celastrina gigas ussuriensis Forster, 1941; Mitt. münchn. ent. Ges. 31 (2): 610; TL: Ussuri, Kassakewitsch
- Celastrina (Celastrina) ladonides; [RFEL]
- Celastrina argiolus heringi; [CoL]
- Celastrina argiolus ladonides; Kudrna, 1974, Atalanta 5: 115; [BRU2], 144; [CoL] ; [iNaturalist]
- Celastrina argiolus levettii; [CoL]
- Celastrina argiolus manchurica; [CoL]
- Celastrina gigas ussuriensis; [CoL]
- Celastrina heringi; [BARU], #361
- Celastrina ladon ladonides; [BARU](note) ,
- Celastrina ladonides ladonides; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 79; [RFEL]
- Celastrina ladonides; [Oleg Kosterin]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 79
- Celastrina sachalinensis; [CoL]
- Cyaniris argiolus ladonides; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 304
- Cyaniris argiolus levetti; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 304
- Lycaenopsis argiolus ladonides; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 28
- Lycaenopsis argiolus levetti; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 28
- ?Lycanopsis ladonides; [BIR], 279 (key), 280 (Murree - Kumaon)
802x642 (~54Kb) underside Deockcheom-ri, Gujwa-eup, Jeju-do Island, South Korea. 31st July 2002, Photo © Oleg Kosterin
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744x570 (~38Kb) underside The Shibu River (Shibukawa) shingle right bank at Hidaka village (Hidakamura), Kochi Prefecture, Shikoku, Japan, 5th August 2002, Photo © Oleg Kosterin
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638x636 (~52Kb) underside The Shibu River (Shibukawa) shingle right bank at Hidaka village (Hidakamura), Kochi Prefecture, Shikoku, Japan, 5th August 2002, Photo © Oleg Kosterin
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Larva on
Prinsepia utilis [BIR],
Lepedeza bicolor,
Vicia amoena,
(Ussuri) [BRU2]: 144 Melilotus ,
Lespedeza ,
Vicia ,
Plectranthus ,
Aralia [RFEL]
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C. a. crimissa (Fruhstorfer, 1917) Formosa
- Celastrina argiolus crimissa; [CoL]
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C. a. canicularis (Verity, 1919) S.Europe
- Celastrina argiolus canicularis; [CoL]
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C. a. grisescens (Eisner, 1964) Netherlands
- argiolus grisescens (Eisner, 1964) (Lycaenopsis); Ent. Ber. 24: 115; TL: Netherlands
- Celastrina argiolus grisescens; [CoL]
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C. a. sugurui Eliot & Kawazoé, 1983 Philippines (Luzon)
- Celastrina argiolus sugurui Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 215, f. 159; TL: Philippines, Luzon, 40km from Baguio, Asin hot spring
Celastrina ladon (Cramer, [1780])
Maryland, West Virginia, .... See [maps]
Spring Azure
- Celastrina argiolus ladon; [BNA], 397; [CoL]
- Celastrina ladon; [NACL], #4363; [NAB], 493; [BARU], #359; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 242; [MPG] ; [P&N23]; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA] ; [GALEPS]
- Celastrina pseudargiolus; [iNaturalist]
- Cyaniris ladon violacea; Dyar, 1903, Bull. U.S. nat. Mus. 52: 45
- Cyaniris ladon; Dyar, 1903, Bull. U.S. nat. Mus. 52: 45; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 304
- Lycaena pseudargiolus; Edwards, 1866, Proc. ent. Soc. Philad. 6 (4): 204 (redescription)
- Lycaena violacea; [LEPIFORUM]
- ?Cyaniris pseudargiolus; [BOW]: pl. 19, f. 11
identification uncertain 600x440 (~43Kb) underside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela
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identification uncertain 800x940 (~96Kb) underside USA: Miller Canyon, Sierra Vista, Cochise Co., Arizona, 8.8.2005, Photo © Markku Savela
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Larva on
Ceanothus ,
Cimicifuga racemosa [NAB],
Malus ,
Physocarpus ,
Prunus ,
Rubus ,
Spiraea ,
Lespedeza ,
Lupinus ,
Melilotus ,
Ledum ,
Arctostaphylos ,
Vaccinium ,
Cimicifuga ,
Lonicera ,
Viburnum ,
Ligustrum ,
Humulus ,
Alnus ,
Cornus ,
Rhus [BARU]